The casparian strip
Casparian strip CS is an impregnation of endodermal cell wall, forming an apoplastic diffusion barrier which forces the symplastic and selective transport of nutrients across endodermis. This extracellular structure can be found in the roots of all higher plants and is thought to provide the protection of vascular tissues. In Arabidopsis , a genetic toolbox regulating the formation of Casparian strips has emerged recently. However, Arabidopsis has the stereotypical root which is much simpler than most other plant species. To understand the Casparian strip formation in a more complex root system, we examined CS regulatory pathways in tomato. Our results reveal a spatiotemporally conserved expression pattern of most essential components of CS machinery in tomato.
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Unveiling the Casparian strip
Unveiling the Casparian strip | Nature
Casparian strip A band of suberin , an impermeable substance, found in the endodermal cell walls of plant roots; it was named after R. Movement of water through the apoplast pathway is diverted from the cell wall to the cytoplasm, where it then follows the symplast pathway. The endodermal cells actively secrete salts into the vascular tissue. This results in a low more negative water potential , allowing water to move down a water potential gradient from the endodermis into the vascular tissue. The Casparian strip prevents water from returning to the cortex; consequently a positive hydrostatic pressure is established in the vascular tissue — the phenomenon of root pressure. January 12, Retrieved January 12, from Encyclopedia.
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In plant anatomy, the Casparian strip named after Robert Caspary is a band of cell wall material deposited in the radial and transverse walls of the endodermis , and is chemically different from the rest of the cell wall - the cell wall being made of lignin  and without suberin - whereas the Casparian strip is made of suberin and sometimes lignin [ clarification needed ]. The chemistry of the Casparian strip has been described as composed of suberin. According to some studies,  the Casparian strip begins as a localized deposition of phenolic and unsaturated fatty substances in the middle lamella between the radial walls, as partly oxidized films.
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